The First Side-Necked Turtle (Pleurodira, Bothremydidae) from the Campanian (Late Cretaceous) of Egypt †

Mohamed AbdelGawad*, Adán Pérez-García, Ren Hirayama, Sara Mohesn, Abdel Aziz Tantawy, Gebely Abu El-Kheir

*Corresponding author for this work

Research output: Contribution to journalArticlepeer-review


The Quseir Formation is an Upper Cretaceous (Campanian) deposit in the Kharga oasis of the Southwestern Desert (Egypt). This formation comprises a clastic sequence of bioturbated mudstone and sandstone intercalations, including rare scattered and fragmented vertebrate remains such as shark teeth, dinosaur remains, and turtle plates. These deposits indicate a supratidal marsh environment. A complete shell of a turtle discovered from the Quseir Formation, at the Kharga oasis, is attributed here to Bothremydini (Pleurodira, Bothremydidae), and determined as a new taxon: Khargachelys caironensis gen. et sp. nov. This form represents the only Bothremydini member currently identified in the Campanian record not only of Egypt but also of North Africa. Therefore, it helps fill the missing evolutionary gap from the Late Cretaceous pleurodires in Egypt and in North Africa.

Original languageEnglish
Article number284
Issue number2
Publication statusPublished - 2023 Feb


  • Bothremydini
  • Campanian
  • Kharga
  • Khargachelys caironensis gen. et sp. nov
  • Quseir Formation

ASJC Scopus subject areas

  • Ecology
  • Ecological Modelling
  • Agricultural and Biological Sciences (miscellaneous)
  • Nature and Landscape Conservation


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